Time delays shape the eco-evolutionary dynamics of cooperation.

We study the intricate interplay between ecological and evolutionary processes through the lens of the prisoner's dilemma game. But while previous studies on cooperation amongst selfish individuals often assume instantaneous interactions, we take into consideration delays to investigate how these might affect the causes underlying prosocial behavior. Through analytical calculations and numerical simulations, we demonstrate that delays can lead to oscillations, and by incorporating also the ecological variable of altruistic free space and the evolutionary strategy of punishment, we explore how these factors impact population and community dynamics. Depending on the parameter values and the initial fraction of each strategy, the studied eco-evolutionary model can mimic a cyclic dominance system and even exhibit chaotic behavior, thereby highlighting the importance of complex dynamics for the effective management and conservation of ecological communities. Our research thus contributes to the broader understanding of group decision-making and the emergence of moral behavior in multidimensional social systems.

Higher-order interactions promote chimera states.

Since the discovery of chimera states, the presence of a nonzero phase lag parameter turns out to be an essential attribute for the emergence of chimeras in a nonlocally coupled identical Kuramoto phase oscillators' network with pairwise interactions. In this Letter, we report the emergence of chimeras without phase lag in a nonlocally coupled identical Kuramoto network owing to the introduction of nonpairwise interactions. The influence of added nonlinearity in the coupled system dynamics in the form of simplicial complexes mitigates the requisite of a nonzero phase lag parameter for the emergence of chimera states. Chimera states stimulated by the reciprocity of the pairwise and nonpairwise interaction strengths and their multistable nature are characterized with appropriate measures and are demonstrated in the parameter spaces.

Spiral wave chimera-like transient dynamics in three-dimensional grid of diffusive ecological systems.

In the present article, we demonstrate the emergence and existence of the spiral wave chimera-like transient pattern in coupled ecological systems, composed of prey-predator patches, where the patches are connected in a three-dimensional medium through local diffusion. We explore the transition scenarios among several collective dynamical behaviors together with transient spiral wave chimera-like states and investigate the long time behavior of these states. The transition from the transient spiral chimera-like pattern to the long time synchronized or desynchronized pattern appears through the deformation of the incoherent region of the spiral core. We discuss the transient dynamics under the influence of the species diffusion at different time instants. By calculating the instantaneous strength of incoherence of the populations, we estimate the duration of the transient dynamics characterized by the persistence of the chimera-like spatial coexistence of coherent and incoherent patterns over the spatial domain. We generalize our observations on the transient dynamics in a three-dimensional grid of diffusive ecological systems by considering two different prey-predator systems.

Amplitude-mediated spiral chimera pattern in a nonlinear reaction-diffusion system.

Formation of diverse patterns in spatially extended reaction-diffusion systems is an important aspect of study that is pertinent to many chemical and biological processes. Of special interest is the peculiar phenomenon of chimera state having spatial coexistence of coherent and incoherent dynamics in a system of identically interacting individuals. In the present article, we report the emergence of various collective dynamical patterns while considering a system of prey-predator dynamics in the presence of a two-dimensional diffusive environment. Particularly, we explore the observance of four distinct categories of spatial arrangements among the species, namely, spiral wave, spiral chimera, completely synchronized oscillations, and oscillation death states in a broad region of the diffusion-driven parameter space. Emergence of amplitude-mediated spiral chimera states displaying drifted amplitudes and phases in the incoherent subpopulation is detected for parameter values beyond both Turing and Hopf bifurcations. Transition scenarios among all these distinguishable patterns are numerically demonstrated for a wide range of the diffusion coefficients which reveal that the chimera states arise during the transition from oscillatory to steady-state dynamics. Furthermore, we characterize the occurrence of each of the recognizable patterns by estimating the strength of incoherent subpopulations in the two-dimensional space.

Persistence in multilayer ecological network consisting of harvested patches.

Complex network theory yields a powerful approach to solve the difficulties arising in a major section of ecological systems, prey-predator interaction being one among them. A large variety of ecological systems have been successfully investigated employing the theory of complex networks, and one of the most significant advancements in this theory is the emerging field of multilayer networks. The field of multilayer networks provides a natural framework to accommodate multiple layers of complexities emerging in ecosystems. In this article, we consider prey-predator patches communicating among themselves while being connected by distinct small-world dispersal topologies in two layers of the network. We scrutinize the robustness of the multilayer ecological network sustaining gradually over harvested patches. We thoroughly report the consequences of introducing asymmetries in both interlayer and intralayer dispersal strengths as well as the network topologies on the global persistence of species in the network. Besides numerical simulation, we analytically derive the critical point up to which the network can sustain species in the network. Apart from the results on a purely multiplex framework, we validate our claims for multilayer formalism in which the patches of the layers are different. Interestingly, we observe that due to the interaction between the two layers, species are recovered in the layer that we assume to be extinct initially. Moreover, we find similar results while considering two completely different prey-predator systems, which eventually attests that the outcomes are not model specific.

Eco-evolutionary dynamics of cooperation in the presence of policing.

Ecology and evolution are inherently linked, and studying a mathematical model that considers both holds promise of insightful discoveries related to the dynamics of cooperation. In the present article, we use the prisoner's dilemma (PD) game as a basis for long-term apprehension of the essential social dilemma related to cooperation among unrelated individuals. We upgrade the contemporary PD game with an inclusion of evolution-induced act of punishment as a third competing strategy in addition to the traditional cooperators and defectors. In a population structure, the abundance of ecologically-viable free space often regulates the reproductive opportunities of the constituents. Hence, additionally, we consider the availability of free space as an ecological footprint, thus arriving at a simple eco-evolutionary model, which displays fascinating complex dynamics. As possible outcomes, we report the individual dominance of cooperators and defectors as well as a plethora of mixed states, where different strategies coexist followed by maintaining the diversity in a socio-ecological framework. These states can either be steady or oscillating, whereby oscillations are sustained by cyclic dominance among different combinations of cooperators, defectors, and punishers. We also observe a novel route to cyclic dominance where cooperators, punishers, and defectors enter a coexistence via an inverse Hopf bifurcation that is followed by an inverse period doubling route.

Cooperation on Interdependent Networks by Means of Migration and Stochastic Imitation.

Evolutionary game theory in the realm of network science appeals to a lot of research communities, as it constitutes a popular theoretical framework for studying the evolution of cooperation in social dilemmas. Recent research has shown that cooperation is markedly more resistant in interdependent networks, where traditional network reciprocity can be further enhanced due to various forms of interdependence between different network layers. However, the role of mobility in interdependent networks is yet to gain its well-deserved attention. Here we consider an interdependent network model, where individuals in each layer follow different evolutionary games, and where each player is considered as a mobile agent that can move locally inside its own layer to improve its fitness. Probabilistically, we also consider an imitation possibility from a neighbor on the other layer. We show that, by considering migration and stochastic imitation, further fascinating gateways to cooperation on interdependent networks can be observed. Notably, cooperation can be promoted on both layers, even if cooperation without interdependence would be improbable on one of the layers due to adverse conditions. Our results provide a rationale for engineering better social systems at the interface of networks and human decision making under testing dilemmas.

Chimera patterns in three-dimensional locally coupled systems.

The coexistence of coherent and incoherent domains, namely the appearance of chimera states, has been studied extensively in many contexts of science and technology since the past decade, though the previous studies are mostly built on the framework of one-dimensional and two-dimensional interaction topologies. Recently, the emergence of such fascinating phenomena has been studied in a three-dimensional (3D) grid formation while considering only the nonlocal interaction. Here we study the emergence and existence of chimera patterns in a three-dimensional network of coupled Stuart-Landau limit-cycle oscillators and Hindmarsh-Rose neuronal oscillators with local (nearest-neighbor) interaction topology. The emergence of different types of spatiotemporal chimera patterns is investigated by taking two distinct nonlinear interaction functions. We provide appropriate analytical explanations in the 3D grid of the network formation and the corresponding numerical justifications are given. We extend our analysis on the basis of the Ott-Antonsen reduction approach in the case of Stuart-Landau oscillators containing infinite numbers of oscillators. Particularly, in the Hindmarsh-Rose neuronal network the existence of nonstationary chimera states is characterized by an instantaneous strength of incoherence and an instantaneous local order parameter. Besides, the condition for achieving exact neuronal synchrony is obtained analytically through a linear stability analysis. The different types of collective dynamics together with chimera states are mapped over a wide range of various parameter spaces.

Resumption of dynamism in damaged networks of coupled oscillators.

Deterioration in dynamical activities may come up naturally or due to environmental influences in a massive portion of biological and physical systems. Such dynamical degradation may have outright effect on the substantive network performance. This requires us to provide some proper prescriptions to overcome undesired circumstances. In this paper, we present a scheme based on external feedback that can efficiently revive dynamism in damaged networks of active and inactive oscillators and thus enhance the network survivability. Both numerical and analytical investigations are performed in order to verify our claim. We also provide a comparative study on the effectiveness of this mechanism for feedbacks to the inactive group or to the active group only. Most importantly, resurrection of dynamical activity is realized even in time-delayed damaged networks, which are considered to be less persistent against deterioration in the form of inactivity in the oscillators. Furthermore, prominence in our approach is substantiated by providing evidence of enhanced network persistence in complex network topologies taking small-world and scale-free architectures, which makes the proposed remedy quite general. Besides the study in the network of Stuart-Landau oscillators, affirmative influence of external feedback has been justified in the network of chaotic Rössler systems as well.

Chimera states in two-dimensional networks of locally coupled oscillators.

Chimera state is defined as a mixed type of collective state in which synchronized and desynchronized subpopulations of a network of coupled oscillators coexist and the appearance of such anomalous behavior has strong connection to diverse neuronal developments. Most of the previous studies on chimera states are not extensively done in two-dimensional ensembles of coupled oscillators by taking neuronal systems with nonlinear coupling function into account while such ensembles of oscillators are more realistic from a neurobiological point of view. In this paper, we report the emergence and existence of chimera states by considering locally coupled two-dimensional networks of identical oscillators where each node is interacting through nonlinear coupling function. This is in contrast with the existence of chimera states in two-dimensional nonlocally coupled oscillators with rectangular kernel in the coupling function. We find that the presence of nonlinearity in the coupling function plays a key role to produce chimera states in two-dimensional locally coupled oscillators. We analytically verify explicitly in the case of a network of coupled Stuart-Landau oscillators in two dimensions that the obtained results using Ott-Antonsen approach and our analytical finding very well matches with the numerical results. Next, we consider another type of important nonlinear coupling function which exists in neuronal systems, namely chemical synaptic function, through which the nearest-neighbor (locally coupled) neurons interact with each other. It is shown that such synaptic interacting function promotes the emergence of chimera states in two-dimensional lattices of locally coupled neuronal oscillators. In numerical simulations, we consider two paradigmatic neuronal oscillators, namely Hindmarsh-Rose neuron model and Rulkov map for each node which exhibit bursting dynamics. By associating various spatiotemporal behaviors and snapshots at particular times, we study the chimera states in detail over a large range of coupling parameter. The existence of chimera states is confirmed by instantaneous angular frequency, order parameter and strength of incoherence.

Survivability of a metapopulation under local extinctions.

A metapopulation structure in landscape ecology comprises a group of interacting spatially separated subpopulations or patches of the same species that may experience several local extinctions. This makes the investigation of survivability (in the form of global oscillation) of a metapopulation on top of diverse dispersal topologies extremely crucial. However, among various dispersal topologies in ecological networks, which one can provide higher metapopulation survivability under local extinction is still not well explored. In this article, we scrutinize the robustness of an ecological network consisting of prey-predator patches having Holling type I functional response, against progressively extinct population patches. We present a comprehensive study on this while considering global, small-world, and scale-free dispersal of the subpopulations. Furthermore, we extend our work in enhancing survivability in the form of sustained global oscillation by introducing asymmetries in the dispersal rates of the considered species. Our findings affirm that the asynchrony among the patches plays an important role in the survivability of a metapopulation. In order to demonstrate the model independence of the observed phenomenon, we perform a similar analysis for patches exhibiting Holling type II functional response. On the grounds of the obtained results, our work is expected to provide a better perception of the influence of dispersal arrangements on the global survivability of ecological networks.